1. State University of New York College of Environmental Science and Forestry, Syracuse, New York13210, United States
2. U.S. Geological Survey, Great Lakes Science Center, Lake Erie Biological Station, 380 Huron Street, Huron, Ohio44839, United States
3. Fairport Fish Research Station, Ohio Department of Natural Resources, 1190 High Street, Fairport Harbor, Ohio44077, United States
4. Department of Aquatic Resources, Swedish University of Agricultural Sciences, SE-750 07Uppsala, Sweden