1. Centre for Cell Engineering, Institute of Molecular, Cell and Systems Biology, College of Medical, Veterinary and Life Sciences, Joseph Black Building, University of Glasgow, Glasgow, G12 8QQ, Scotland, U.K.
2. Glasgow Polyomics Facility, Joseph Black Building, Institute of Biomedical and Life Sciences, University of Glasgow, Glasgow, G12 8QQ, Scotland, U.K.
3. Division of Biomedical Engineering, School of Engineering, University of Glasgow, Glasgow, G12 8LT, Scotland, U.K.
4. Bone & Joint Research Group, Centre for Human Development, Stem Cells and Regeneration, Institute of Developmental Sciences, University of Southampton, Southampton SO16 6YD, U.K.