1. Institute of Coastal Environmental Pollution Control, Ministry of Education Key Laboratory of Marine Environment and Ecology, College of Environmental Science and Engineering, Frontiers Science Center for Deep Ocean Multispheres and Earth System, Ocean University of China, Qingdao 266100, China
2. Marine Ecology and Environmental Science Laboratory, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266071, China
3. Ministry of Ecology and Environment, South China Institute of Environmental Sciences, Guangzhou 510535, China
4. Sanya Oceanographic Institution, Ocean University of China, Sanya 572000, China
5. Stockbridge School of Agriculture, University of Massachusetts, Amherst, Massachusetts 01003, United States