1. Institute of Grassland and Environmental Research, North Wyke, Okehampton, Devon EX20 2SB, United Kingdom
2. Department of Agricultural and Environmental Science, University of Newcastle upon Tyne, Newcastle upon Tyne NE1 7RU, United Kingdom
3. German Institute of Human Nutrition, Potsdam-Rehbrucke, Bergholz-Rehbrucke, D-14858, Germany
4. Institute of Grassland and Environmental Research, Plas Gogerddan, Aberystwyth, Ceredigion SY23 3EB, United Kingdom