1. JST CREST, Sanbancho-bldg., 5 Sanbancho, Chiyoda-ku, Tokyo, 102-0075, Japan
2. Institut des Sciences et Ingénierie Chimiques, Ecole Polytechnique Fédérale de Lausanne, CH-1015, Lausanne, Switzerland
3. Department of Cell Biology, Center for Cell Dynamics, School of Medicine, Johns Hopkins University, Baltimore, Maryland 21205, United States