1. Food Research Program, Agriculture and Agri-Food Canada, Guelph, Ontario, Canada; Crop Development Centre and Department of Plant Sciences and Department of Biology, University of Saskatchewan, Saskatoon, Saskatchewan, Canada; Lethbridge Research Centre, Agriculture and Agri-Food Canada, Lethbridge, Alberta, Canada; and Eastern Cereal and Oilseed Research Centre, Agriculture and Agri-Food Canada, Ottawa, Ontario, Canada