1. Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences, Beijing, 100085, China. Institute of Biological and Environmental Sciences, University of Aberdeen, Cruickshank Building, St. Machar Drive, Aberdeen, AB24 3UU, U.K., Institute of Urban Environment, Chinese Academy of Sciences, Xiamen, 361003, China, and College of Natural Resources and Environment, South China Agricultural University, Guangzhou, 510642 China