1. Department of Molecular Genetics, University of Toronto, Toronto, Ontario M5S 1A8, Canada
2. Department of Biochemistry, University of Toronto, Toronto, Ontario M5S 1A8, Canada
3. Department of Chemistry, University of Toronto, Toronto, Ontario M5S 1A8, Canada
4. Hospital for Sick Children, Program in Molecular Medicine, 555 University Avenue, Toronto, Ontario M5G 1X8, Canada