1. Department of Biosciences, University of Kuopio, P.O. Box 1627, FIN-70211 Kuopio, Finland; Department of Biology, University of Fribourg, CH-1700 Fribourg, Switzerland; Quality Health and Nutrition Department, Scottish Crop Research Institute, Invergowrie, Dundee, DD2 5DA, United Kingdom; and MTT Agrifood Research Finland, FIN-31600 Jokioinen, Finland