1. Program in Chemical Biology, Department of Physiology and Pharmacology, Oregon Health & Science University, 3181 SW Sam Jackson Park Road, Portland, Oregon 97239, United States
2. Department of Embryology, Carnegie Institution for Science, 3520 San Martin Drive, Baltimore, Maryland 21218, United States
3. Knight Cancer Institute, Oregon Health & Science University, 3181 SW Sam Jackson Park Road, Portland, Oregon 97239, United States