1. Key Laboratory of Tropical Marine Bio-resources and Ecology, Guangdong Key Laboratory of Marine Materia Medica, Innovation Academy of South China Sea Ecology and Environmental Engineering, South China Sea Institute of Oceanology, Chinese Academy of Sciences, 164 West Xingang Road, Guangzhou 510301, China
2. University of Chinese Academy of Science, 19 Yuquan Road, Beijing 100049, China
3. Sanya Institute of Oceanology Eco-Environmental Engineering, Yazhou Scientific Bay, Sanya 572000, China