1. Faculty and Graduate School of Fisheries Sciences, Hokkaido University, 3-1-1 minato-cho, Hakodate, Hokkaido 041-8611, Japan
2. Frontier Research Center for Advanced Material and Life Science, Faculty of Advanced Life Science, Hokkaido University, Kita 21 Nishi 11, Sapporo 001-0021, Japan
3. University of the Ryukyus Senbaru 1, Nishihara, Okinawa 903-0213, Japan