1. Academic Unit of Reproductive and Developmental Medicine, University of Sheffield, Level 4, The Jessop Wing, Tree Root Walk, Sheffield S10 2SF, United Kingdom, Biological and Environmental Systems Group, Department of Chemical and Process Engineering, University of Sheffield, Mappin Street, Sheffield S1 3JD, United Kingdom, and Department of Medicine and Animal Surgery, Faculty of Veterinary Medicine, University of Murcia, 30.071 Murcia, Spain