1. Department of Microbiology, Faculty of Science, Helwan University, Ain Helwan, Helwan, Cairo 11795, Egypt
2. Department of Biotechnology, Delft University of Technology, van der Maasweg 9, 2629 HZ Delft, The Netherlands
3. Laboratory of Organic Chemistry, Wageningen University & Research, Stippeneng 4, 6708 WE Wageningen, The Netherlands