1. School of Bioscience and Bioengineering, South China University of Technology, Guangzhou 510006, PR China
2. School of Food Sciences and Engineering, South China University of Technology, Guangzhou 510640, PR China
3. Department of Biotechnology, Delft University of Technology, Van der Maasweg 9, 2629HZ Delft, The Netherlands
4. Institute of Biochemistry, Department of Biotechnology and Enzyme Catalysis, Greifswald University, Felix-Hausdorff-Str. 4, 17487 Greifswald, Germany