1. Key Laboratory of Industrial Fermentation Microbiology of the Ministry of Education, Tianjin Key Laboratory of Industrial Microbiology, College of Biotechnology, Tianjin University of Science and Technology, National Engineering Laboratory for Industrial Enzymes, Tianjin 300457, People’s Republic of China
2. College of Bioengineering, Qilu University of Technology, Jinan 250100, People’s Republic of China