1. Department of Environmental Chemistry, IIQAB-CSIC. Jordi Girona 18-26, 08034 Barcelona, Catalonia, Spain; Aquatic Biotechnology, Biofilm Center, University Duisburg-Essen, Geibelstr. 41, 47057 Duisburg, Germany; Department of Environmental Microbiology, Helmholtz Centre for Environmental Research − UFZ. Permoserstrasse 15, 04318 Leipzig, Germany; Department of Isotope Biogeochemistry; Helmholtz Centre for Environmental Research − UFZ. Permoserstrasse 15, 04318 Leipzig, Germany