1. Marine Life Support
Systems, Plymouth Marine Laboratory, Prospect Place, The Hoe, Plymouth PL1 3DH, U.K.
2. College
of Life, Environmental Sciences: Biosciences, Geoffrey Pope Building, University of Exeter, Stocker Road, Exeter EX4
4QD, U.K.
3. Akvaplan-niva AS,
FRAM − High North Research Centre for Climate and the Environment, N-9296 Tromsø, Norway