1. Agricultural College, Shenyang Agricultural University, Shenyang 110866, PR China
2. College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, PR China
3. Cardiff University, School of Biosciences, Cardiff CF10 3TL, U.K.
4. Key Laboratory of Pollution Ecology and Environmental Engineering, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, PR China