1. Engineering Research Center of Industrial Microbiology, Ministry of Education; Collaborative Innovation Center of Haixi Green Bio-Manufacturing Technology, Ministry of Education; National & Local Joint Engineering Research Center of Industrial Microbiology and Fermentation Technology, National Development and Reform Commission; College of Life Sciences, Fujian Normal University, Fuzhou, 350117, P. R. China
2. CAS Key Laboratory of Special Pathogens and Biosafety, Wuhan Institute of Virology, Wuhan 430071, P. R. China
3. The State Key Laboratory of Virology, Wuhan 430071, P. R. China