1. Sanya Nanfan Research Institute of Hainan University, Sanya 572025, China
2. Hainan Yazhou Bay Seed Laboratory, Sanya 572025, China
3. College of Tropical Crops, Hainan University, Haikou 570288, China
4. College of Life Sciences, Jilin Agricultural University, Changchun 130118, China
5. School of Chemistry and Chemical Engineering, Guangzhou University, Guangzhou 510006, China