1. College of Tropical Crops, Hainan University, Haikou 570228, China
2. State Key Laboratory of Hulless Barley and Yak Germplasm Resources and Genetic Improvement, Lhasa 850002, China
3. Agricultural Research Institute, Tibet Academy of Agricultural and Animal Husbandry Sciences Lhasa, Tibet 850002, China
4. Plant Sciences College, Tibet Agriculture & Animal Husbandry University, Nyingchi, Tibet 860000, China
5. Wuhan Metware Biotechnology Co., Ltd., Wuhan 430070, China