1. State Key Laboratory of Soil and Sustainable Agriculture, Institute of Soil Science, Chinese Academy of Sciences, Nanjing 210008, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. College of Resource and Environment, Anhui Science and Technology University, Chuzhou 233100, China
4. Northeast Institute of Geography and Agricultural Ecology, Chinese Academy of Sciences, Harbin 150040, China
5. School of Biological Sciences, University of Oklahoma, Norman, Oklahoma 73069, United States