1. College
of Bee Sciences/College of Life Sciences, Fujian Agriculture and Forestry University, Fuzhou, Fujian 350002, People’s Republic of China
2. Institute
of Apicultural Research, Chinese Academy of Agricultural Sciences, Beijing 100093, People’s Republic of China
3. Bee
Research Laboratory, Agricultural Research Service (ARS), United States Department of Agriculture (USDA), Beltsville, Maryland 20705, United States