1. Key Laboratory of Saline-alkali Vegetation Ecology Restoration (Northeast Forestry University), Ministry of Education, Harbin 150040, China
2. College of Life Science, Northeast Forestry University, Harbin 150040, China
3. Heilongjiang Key Laboratory of Plant Bioactive Substance Biosynthesis and Utilization, Northeast Forestry University, Harbin 150040, China
4. Yunnan Enov Bioengineering Co., Ltd, 2nd Floor, Building B2, 16 PuFa Road, Export Processing Zone, Economic Development Zone, Kunming, Yunnan 650217, China
5. Department of Forest Resources, College of Forest and Environmental Sciences, Kangwon National University, Chunchon 200-701, Republic of Korea