1. School of Marine Sciences, Sun Yat-sen University, Zhuhai 519082, P. R. China
2. Key Laboratory for Biobased Materials and Energy of Ministry of Education, College of Materials and Energy, South China Agricultural University, Guangzhou, 510642, China
3. Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, Zhuhai, China, 519082
4. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519000, P. R. China
5. Pearl River Estuary Marine Ecosystem Research Station, Ministry of Education, Zhuhai, 519082, China