1. CAS Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, Shandong 266101, China
2. Shandong Provincial Key Laboratory of Energy Genetics, Shandong Engineering Laboratory of Single Cell Oil, Shandong Energy Institute, Qingdao, Shandong 266101, China
3. Qingdao Engineering Laboratory of Single Cell Oil, Qingdao New Energy Shandong Laboratory, Qingdao, Shandong 266101, China
4. Academy of Plateau Science and Sustainability, Qinghai Normal University, Xining, Qinghai 810016, China