1. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations, CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, People’s Republic of China
2. University of Chinese Academy of Sciences, Beijing 100049, People’s Republic of China
3. State Key Laboratory of Phytochemistry and Plant Resources in West China, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, People’s Republic of China
4. Guizhou University of Traditional Chinese Medicine, Guiyang, Guizhou 550002, People’s Republic of China