1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao 266237, China
2. College of Bioengineering, Qilu University of Technology, Jinan 250100, China
3. Yantai Huakangrongzan Biotechnology Co., Ltd, Yantai 264006, China
4. Key Laboratory of Systems Microbial Biotechnology, Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, China
5. National Glycoengineering Research Center, Shandong University, Qingdao, Shandong 266237, China