1. State Key Laboratory for Conservation and Utilization of Subtropical Agro-bioresources, South China Agricultural University, Guangzhou 510642, China
2. Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant Germplasm, Guangzhou 510642, China
3. Mid-Florida Research & Education Center, IFAS, University of Florida, Apopka, Florida 32703, United States
4. College of Horticulture, Hebei Agricultural University, Hebei 071066, China