1. State Key Laboratory of Microbial Technology, Shandong University, No. 72 Binhai Road, Qingdao, Shandong 266237, China
2. Angel Yeast Co., Ltd., Yichang, Hubei 443003, China
3. National Key Laboratory of Agricultural Microbiology, Yichang, Hubei 443003, China
4. Laboratory for Marine Biology and Biotechnology, Qingdao Marine Science and Technology Center, Qingdao, Shandong 266237, China