1. Faculty of Sciences, Centre for Environmental Biology (CBA), University of Lisbon, FCUL, Campo Grande, Bloco C2, Piso 5, 1749-016 Lisboa, Portugal, Portuguese Environmental Protection Agency (APA), Rua da Murgueira, 9/9a, 2611-865 Amadora, Portugal, and Centre for Natural Resources and the Environment (CERENA), IST, UTL, Av. Rovisco Pais, 1049-001 Lisboa, Portugal