1. Department of Chemistry and Biotechnology, School of Engineering, University of Tokyo, Hongo, Bunkyo-ku, Tokyo 113-8656, Japan, Department of Integrated Biosciences, Graduate School of Frontier Sciences, University of Tokyo, Building FSB-401, 5-1-5 Kashiwanoha Kashiwa Chiba 277-8562, Japan, and Research Center for Environmental Genomics, Kobe University, 1-1 Rokkodai-cho, Nada-ku, Kobe 657-8501, Japan