1. Key Laboratory of Brain Function and Diseases of Chinese Academy of Sciences, School of Life Sciences, University of Science and Technology of China, Hefei, Anhui 230027, P. R. China, Hefei National Laboratory for Physical Sciences at the Microscale, University of Science and Technology of China, Hefei, Anhui 230027, P. R. China, and Suzhou GenePharma Co., Ltd, Suzhou, Jiangsu 215123, P. R. China