1. Department of Agricultural and Biosystems Engineering, Macdonald Campus, McGill University, 21111 Lakeshore Road, Ste. Anne-de-Bellvue, Quebec, H9X 3V9 Canada, and Centre de Microbiologie et Biotechnologie, INRS-Institut Armand-Frappier, 245 Boulevard Hymus, Pointe-Claire, Quebec, H9R 1G6 Canada