1. ATTA. 2004. Sistema de información de INBio, Animalia. http://www.inbio.ac.cr/bims/k02/p05/c029.htm .
2. Burns, J.M., and D.H. Janzen. 2001. Biodiversity of pyrrhopygine skipper butterfies (Hesperiidae) in the Area de Conservación Guanacaste, Costa Rica. Journal of the Lepidopterists' Society 55: 15–43.
3. Cohen, A.C., and J.D. Pinto. 1977. An evaluation of xeric adaptiveness of several species of blister beetles (Meloidae). Annals of the Entomological Society of America 70: 741–49.
4. DeVries, P.J. 1987. The Butterfies of Costa Rica and Their Natural History: Papilionidae, Pieridae, Nymphalidae. Princeton, NJ: Princeton University Press.
5. ——. 1997. The Butterfies of Costa Rica and Their Natural History. Vol. 2. Riodinidae. Princeton, NJ: Princeton University Press.