1. Institute of Earth Sciences (ISTE), University of Lausanne, Geopolis, CH-1015 Lausanne, Switzerland;
2. Institute of Earth Sciences (ISTE), University of Lausanne, Geopolis, CH-1015 Lausanne, Switzerland,
3. Spatial Ecology and Paleontology laboratory (SEPL), School of Biological Sciences, Georgia Institute of Technology, Atlanta, Georgia, USA;