1. Department of Zoology, University of Oxford, Zoology Research and Administration Building, 11a Mansfield Road, Oxford OX1 3SZ, UK
2. Imperial College London, Exhibition Road, South Kensington, London SW7 2BX, UK
3. Barnside, Buckland Brewer, Bideford, Devon EX39 5NF, UK
4. Oxford Martin School, University of Oxford, 34 Broad Street, Oxford OX1 3BD, UK