1. Odum School of Ecology, College of Veterinary Medicine, University of Georgia, Athens, GA, USA
2. Center for the Ecology of Infectious Disease, College of Veterinary Medicine, University of Georgia, Athens, GA, USA
3. Department of Biology, Emory University, Atlanta, GA, USA
4. Wisconsin Department of Natural Resources, Madison, WI, USA
5. Department of Infectious Diseases, College of Veterinary Medicine, University of Georgia, Athens, GA, USA