1. Evolutionary Ecology of Variation Research Group, Max Planck Institute for Ornithology, Eberhard-Gwinner-Straße 7, Seewiesen (Starnberg) 82319, Germany
2. Department of Behavioural Ecology and Evolutionary Genetics, Max Planck Institute for Ornithology, Eberhard-Gwinner-Straße 7, Seewiesen (Starnberg) 82319, Germany
3. Behavioural Ecology, Department Biologie der LMU, Ludwig Maximilians University of Munich (LMU), Großhadener Straße. 2, Planegg-Martinsried 82152, Germany