1. Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA 02138, USA
2. CNRS, Université Paul Sabatier, ENFA, UMR 5174 EDB (Laboratoire Évolution et Diversité Biologique), Université de Toulouse, 118 route de Narbonne, Toulouse 31062, France
3. Department of Botany and Zoology, University of Stellenbosch, Private Bag X1, Matieland 7602, South Africa
4. Landcare Research, Auckland Mail Centre, Private Bag 92170, Auckland 1142, New Zealand
5. School of Biological Sciences, University of Auckland, Auckland, New Zealand
6. Allan Wilson Centre for Molecular Ecology and Evolution, Auckland, New Zealand
7. Animal Evolution and Development, Institute of Biology, University of Leipzig, Talstraße 33, Leipzig 04103, Germany