1. School of Ocean and Earth Science, National Oceanography Centre Southampton, University of Southampton, European Way, Southampton SO14 3ZH, UK
2. School of Ocean Sciences, Bangor University, Bangor LL57 2DG, UK
3. Centre for Environment, Fisheries and Aquaculture Science, Lowestoft Laboratory, Pakefield Road, Lowestoft, Suffolk NR33 0HT, UK