1. Palau International Coral Reef Center, PO Box 7086, Koror, Palau
2. Marine Ecology Research Centre, Southern Cross University, PO Box 157, Lismore, New South Wales 2480, Australia
3. Australian Institute of Marine Science, PMB 3, Townsville, Queensland 4810, Australia
4. TropWATER and College of Marine and Environmental Sciences, James Cook University, Townsville, Queensland, Australia
5. CSIRO Oceans and Atmosphere, St Lucia, Queensland 4067, Australia