1. State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, People's Republic of China
2. Graduate University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China
3. UMR 7625, UPMC-CNRS-ENS, Ecole Normale Superieure, 46 rue d'Ulm, 75230 Paris cedex 05, France
4. UMI 209, IRD-UPMC, 32 avenue Henri Varagnat, 93142 Bondy cedex, France
5. Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biology, University of Oslo, PO Box 1066, Blindern 0316, Oslo, Norway
6. Friedrich-Alexander University Erlangen-Nuremberg, Institute of Geography, Kochstrasse 4/4, 91054 Erlangen, Germany
7. Institute of World History, The Chinese Academy of Social Sciences, Beijing 100732, People's Republic of China