1. Department of Biology, University of Utah, Salt Lake City, UT 84112, USA
2. Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA
3. Department of Entomology, University of Maryland, College Park, MD 20742, USA
4. Center for Social Insect Research, School of Life Sciences, Arizona State University, Tempe, AZ 85287, USA
5. Department of Biological Sciences and Museum of Natural Science, Louisiana State University, Baton Rouge, LA 70803, USA