1. Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY 14853, USA
2. Oregon Institute of Marine Biology, University of Oregon, Charleston, OR 97420, USA
3. Marine Geophysical Laboratory, Physics Department, College of Science and Engineering, James Cook University, Townsville, Queensland 4811, Australia
4. NOAA Coral Reef Watch, NESDIS Center for Satellite Applications and Research, College Park, MD 20740, USA