1. Department of Ecology and Evolution, University of Lausanne, 1015 Lausanne, Switzerland
2. Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, University of Glasgow, Graham Kerr Building, Glasgow G12 8QQ, UK
3. Cardiff University School of Biosciences, Biomedical Sciences Building, Museum Avenue, Cardiff CF10 3AX, UK