1. Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA 02138, USA
2. Department of Biology, University of Florida, 876 Newell Drive, Gainesville, FL 32611, USA
3. Department of Life Sciences, Imperial College London, Silwood Park Campus, Buckhurst Road, Ascot, Berkshire SL5 7PY, UK
4. Natural History Museum, Cromwell Road, London SW7 5BD, UK