1. School of Mathematics and Statistics, College of Science and Engineering, Institute of Infection, Immunity and Inflammation, University of Glasgow, Glasgow, UK
2. The Boyd Orr Centre for Population and Ecosystem Health, Institute of Infection, Immunity and Inflammation, University of Glasgow, Glasgow, UK
3. Institute of Biodiversity, Animal Health and Comparative Medicine, College of Medical, Veterinary and Life Sciences, Institute of Infection, Immunity and Inflammation, University of Glasgow, Glasgow, UK
4. Wellcome Trust Centre for Molecular Parasitology, Institute of Infection, Immunity and Inflammation, University of Glasgow, Glasgow, UK