1. Evolutionary Biology, Institute of Biology Leiden, Leiden University, RA 2300 Leiden, The Netherlands
2. University Museum of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
3. Department of Analytical Chemistry, Gembloux Agro-Bio Tech, Liège University, Passage des Déportés 2, 5030 Gembloux, Belgium
4. Laboratory of Genetics, Wageningen University, BD 6703 Wageningen, The Netherlands
5. Evolutionary Ecology and Genetics Group, Biodiversity Research Centre, Earth and Life Institute, Académie Louvain, Croix du Sud 4, 1348 Louvain-la-Neuve, Belgium